KEGG   ENZYME: 3.2.1.4Help
Entry
EC 3.2.1.4                  Enzyme                                 

Name cellulase;
endo-1,4-beta-D-glucanase;
beta-1,4-glucanase;
beta-1,4-endoglucan hydrolase;
celluase A;
cellulosin AP;
endoglucanase D;
alkali cellulase;
cellulase A 3;
celludextrinase;
9.5 cellulase;
avicelase;
pancellase SS;
1,4-(1,3;
1,4)-beta-D-glucan 4-glucanohydrolase
Class Hydrolases;
Glycosylases;
Glycosidases, i.e. enzymes that hydrolyse O- and S-glycosyl
compounds
BRITE hierarchy
Sysname 4-beta-D-glucan 4-glucanohydrolase
Reaction(IUBMB) Endohydrolysis of (1->4)-beta-D-glucosidic linkages in cellulose,
lichenin and cereal beta-D-glucans
Reaction(KEGG) (other) R02886 R06200(G)
Show
Comment Will also hydrolyse 1,4-linkages in beta-D-glucans also containing
1,3-linkages.
Pathway PATH: ec00500  Starch and sucrose metabolism
Orthology KO: K01179  endoglucanase
Genes ATH: AT3G43860 AT4G39000 AT4G39010
OSA: 4330912(Os02g0778600) 4334029(Os03g0736300)
     4335944(Os04g0443300) 4347660(Os09g0530200)
ANI: AN1285.2
AFM: AFUA_3G03870 AFUA_5G01830 AFUA_6G01800 AFUA_6G07480
     AFUA_7G01540 AFUA_7G06740
AOR: AO090003001342
ANG: An07g08950(eglB) An14g02760(eglA)
CNE: CNH00790
DDI: DDB_0215351(celA) DDB_0230001
ECO: b3531(bcsZ)
ECJ: JW3499(bcsZ)
ECD: ECDH10B_3708(bcsZ)
EBW: BWG_3220(bcsZ)
ECE: Z4946(yhjM)
ECS: ECs4411
ECF: ECH74115_4896(bcsZ)
ETW: ECSP_4521(bcsZ)
ECG: E2348C_3773(bcsZ)
ECC: c4343(yhjM)
ECI: UTI89_C4063(yhjM)
ECP: ECP_3631
ECV: APECO1_2917(bcsZ)
ECW: EcE24377A_4019(bcsZ)
ECM: EcSMS35_3840(bcsZ)
ECY: ECSE_3800
ECL: EcolC_0186
ECK: EC55989_3978(bcsZ)
ECQ: ECED1_4209(bcsZ)
ECR: ECIAI1_3682(bcsZ)
ECT: ECIAI39_4034(bcsZ)
ECZ: ECS88_3945(bcsZ)
EUM: ECUMN_4039(bcsZ)
ELF: LF82_0217(bcsZ)
EBL: B21_03332(bcsZ)
EBD: ECBD_0208
EBR: ECB_03379(bcsZ)
EFE: EFER_3516(bcsZ)
STY: STY4183(yhjM)
STT: t3900(yhjM)
SPT: SPA3473(yhjM)
SEK: SSPA3243
SPQ: SPAB_04494
SEI: SPC_3692(yhjM)
SEC: SC3551
SEH: SeHA_C3933(bcsZ)
SEE: SNSL254_A3889(bcsZ)
SEW: SeSA_A3812(bcsZ)
SEA: SeAg_B3825(bcsZ)
SED: SeD_A3993(bcsZ)
SEG: SG3819(bcsZ)
SET: SEN3440(bcsZ)
SES: SARI_04017
STM: STM3617
YPK: y3831
YPA: YPA_3825
YPN: YPN_3647
YPP: YPDSF_3364
YPS: YPTB3837
YPI: YpsIP31758_4070(bcsZ)
YPY: YPK_0093
YPB: YPTS_4053
YEN: YE4072A(bcsZ)
SFX: S4204(yhjM)
SFV: SFV_3557(yhjM)
SSN: SSON_3860(yhjM)
SBO: SBO_3530(yhjM)
SBC: SbBS512_E3937(bcsZ)
ECA: ECA1981(celV) ECA4373(bcsZ)
ETA: ETA_33880(celY)
ENT: Ent638_3928 Ent638_3936
ESA: ESA_04206
KPN: KPN_03881(bcsC) KPN_03889
KPE: KPK_0216 KPK_0224(bcsZ)
KPU: KP1_5222(bcsC) KP1_5232
CKO: CKO_04974
SPE: Spro_0150
PMR: PMI3103(celY)
DDA: Dd703_3891
DZE: Dd1591_4079
XFA: XF0818 XF2708
XFT: PD1851(engXCA) PD2061(egl)
XFM: Xfasm12_2256
XFN: XfasM23_1953 XfasM23_2164
XCC: XCC0026(egl) XCC0027(egl) XCC0028(egl) XCC2387(egl2)
     XCC3521(engXCA)
XCB: XC_0026 XC_0027 XC_0028 XC_0639 XC_1727
XCA: xccb100_0029(egl1) xccb100_0030(egl2) xccb100_0031(egl3)
     xccb100_0675(engXCA)
XCV: XCV0029(egl1) XCV0031(egl2) XCV0033(egl3) XCV0358
     XCV0670(engXCA) XCV2704(egl4) XCV3641(bcsZ)
XAC: XAC0028(egl) XAC0029(egl) XAC0030(egl) XAC0612(engXCA)
     XAC2522(egl2) XAC3516
XOO: XOO0281(egl) XOO0282(egl) XOO0283(egl) XOO4019(engXCA)
XOM: XOO_0253 XOO_0254 XOO_0255 XOO_3789
VFI: VF_A0882(bcsZ)
VFM: VFMJ11_A1001
PPR: PBPRA1722 PBPRB0304
PPU: PP_2637
PPF: Pput_2138
PPG: PputGB1_3195
PST: PSPTO_1029(wssD)
CJA: CJA_0619(cel5H) CJA_1633(cel9B) CJA_2983(cel5B) CJA_3337(cel5E)
PHA: PSHAa2164(bcsZ)
PAT: Patl_1404
SDE: Sde_0636 Sde_2490 Sde_2636 Sde_3237 Sde_3239
CVI: CV_2676(bscZ)
RSO: RSp0162(egl)
REU: Reut_B3526
RME: Rmet_2251
CTI: RALTA_B0291(bcsZ)
BMA: BMAA1589(bcsC)
BML: BMA10229_2008(bcsC)
BMN: BMA10247_A0686(bcsC)
BPS: BPSS1581(bcsZ)
BPM: BURPS1710b_A0632(bcsZ)
BPL: BURPS1106A_A2145
BPD: BURPS668_A2231
BTE: BTH_II0792
BVI: Bcep1808_1350
BUR: Bcep18194_A4526
BCN: Bcen_0898
BCH: Bcen2424_1380
BCM: Bcenmc03_1358
BCJ: BCAL1390(bcsZ)
BAM: Bamb_1259
BAC: BamMC406_1292
BMU: Bmul_1925
BMJ: BMULJ_01315(egl)
BXE: Bxe_B2042
BPH: Bphy_2378 Bphy_3254
BPY: Bphyt_5838
PNU: Pnuc_1167
BAV: BAV2628(bcsZ)
AAV: Aave_2102
LCH: Lcho_2071
AZO: azo2236(eglA)
GSU: GSU2196
GME: Gmet_2294
GUR: Gura_3125
PCA: Pcar_1216(sgcX)
MXA: MXAN_4837(celA)
SCL: sce1468(celA) sce4910(celB)
SMD: Smed_5210
RET: RHE_CH00072(ypch00021) RHE_CH00228(egl) RHE_CH01544(celC)
REC: RHECIAT_CH0000271 RHECIAT_CH0001612(celC)
RLE: RL0081 RL0236 RL1648(celY)
RLT: Rleg2_1206 Rleg2_4209
BME: BMEII0724
BJA: blr3367
RPA: RPA4584
MEX: Mext_0058 Mext_1369
MDI: METDI2019
MRD: Mrad2831_0273 Mrad2831_4797
MET: M446_0105 M446_1066
MPO: Mpop_1314
MCH: Mchl_1569 Mchl_4803
MNO: Mnod_3739
RSP: RSP_0157
RSH: Rsph17029_1976
RSQ: Rsph17025_0964
MMR: Mmar10_2725
ZMO: ZMO1086
ACR: Acry_0570
GDI: GDI_2537
GDJ: Gdia_0779
BSU: BSU18130(eglS)
BHA: BH0603 BH2105
BCZ: BCZK2421(choA) pE33L466_0086(celCCC)
BLI: BL01471(bglC)
BLD: BLi01880 BLi01882 BLi02088(bglC)
BCL: ABC3362
BAY: RBAM_018100(bglC)
OIH: OB2238 OB2332
SAB: SAB1605c SAB2338c
SAJ: SaurJH9_2482
SAH: SaurJH1_2531
LLM: llmg_1166 llmg_1167
CAC: CAC0214 CAC0215
CNO: NT01CX_0791(celM) NT01CX_0793
CTH: Cthe_0040 Cthe_0607 Cthe_0608 Cthe_0609 Cthe_0825
CDF: CD2546
AMT: Amet_2620 Amet_2621
STH: STH1079 STH1080 STH1081
ATE: Athe_0594
DSY: DSY1620 DSY1621 DSY1622
EEL: EUBELI_20587
ERE: EUBREC_0616
TTE: TTE0359 TTE0441(frvX) TTE0442(frvX2) TTE0443(frvX3)
TEX: Teth514_0377
TPD: Teth39_1874
CSC: Csac_0678 Csac_1079
NTH: Nther_0313
UUR: UU280(celM)
AYW: AYWB_658(ysdC)
MTU: Rv0062(celA1) Rv1089A(celA2a) Rv1090(celA2b)
MTC: MT0067(celA)
MRA: MRA_0064(celA1) MRA_1100(celA2a) MRA_1101(celA2b)
MTB: TBMG_00061(TBMG_00061.1)
MBO: Mb0063(celA1) Mb1119(celA2a) Mb1120(celA2b)
MBB: BCG_0093(celA1) BCG_1149(celA2a) BCG_1150(celA2b)
MBT: JTY_0063(celA1)
MAV: MAV_0326
MSM: MSMEG_6752
MUL: MUL_0371
MVA: Mvan_3505 Mvan_5837
MGI: Mflv_1003 Mflv_3001
MAB: MAB_4777c
MMC: Mmcs_5301
MKM: Mkms_5390
MJL: Mjls_5680
CMI: CMM_2443(celB) pCM1_0020(celA)
NCA: Noca_3501 Noca_4617
TFU: Tfu_0901 Tfu_1074 Tfu_1627 Tfu_2176 Tfu_2712
FRE: Franean1_7197
FAL: FRAAL4955(celA1) FRAAL4956(celA2)
ACE: Acel_0135
SEN: SACE_4004(bglC) SACE_6060(celA)
SUS: Acid_2606 Acid_4467
CHU: CHU_1401(cel) CHU_1655(cel) CHU_1842(cel) CHU_2103(cel)
     CHU_2802(bglC)
CPI: Cpin_6252
FJO: Fjoh_4946
COC: Coch_0518
AMU: Amuc_0867
RBA: RB5256 RB8541(celM)
SYN: slr0897
SYG: sync_1467 sync_2705
CCH: Cag_0339
PLT: Plut_0993
TRO: trd_A0877
DRA: DR_0229
DGE: Dgeo_1994
AAE: aq_1401(celY)
SAF: SULAZ_1375
TMA: TM1048 TM1049 TM1050 TM1524 TM1751 TM1752
TPT: Tpet_1701 Tpet_1702 Tpet_1703
TLE: Tlet_1475
TRQ: TRQ2_1758 TRQ2_1759
TNA: CTN_1517 CTN_1518
TNP: Tnap_1715 Tnap_1716
TME: Tmel_0097 Tmel_0208
TAF: THA_329 THA_442
FNO: Fnod_1077
PMO: Pmob_1120 Pmob_1121
KOL: Kole_0038
DTH: DICTH_0008 DICTH_0505 DICTH_0508 DICTH_0516 DICTH_0517
     DICTH_1476
DTU: Dtur_0276 Dtur_0669 Dtur_0671 Dtur_0680 Dtur_0681 Dtur_1586
MJA: MJ0555
MMP: MMP1116
MMQ: MmarC5_0468
MMX: MmarC6_1551
MMZ: MmarC7_0368
MAE: Maeo_0709
MVN: Mevan_0441
MAC: MA3849
MBA: Mbar_A0214
MMA: MM_0673
MBU: Mbur_0712
MEM: Memar_1505
MBN: Mboo_1201
MPL: Mpal_2409
MSI: Msm_0134
MKA: MK0383
AFU: AF1795(celM)
HAL: VNG1498G(celM)
HMA: rrnAC0799(cdlM)
HWA: HQ2923A(celM)
NPH: NP4306A(celM)
HLA: Hlac_0724
PHO: PH1171 PH1527
PAB: PAB0437 PAB0632(celB-like)
PFU: PF1547
TKO: TK0781
TON: TON_1032
TSI: TSIB_1215
SMR: Smar_0057
DKA: DKAM_0589
HBU: Hbut_1154
PAI: PAE1385
PIS: Pisl_1432
PCL: Pcal_0842
PAS: Pars_0452
CMA: Cmaq_0206 Cmaq_0950
TNE: Tneu_0542
TPE: Tpen_0002 Tpen_0177
KCR: Kcr_0883
Taxonomy
Structures PDB: 1A39  1A3H  1AIW  1CEC  1CEM  1CEN  1CEO  1CLC  1CX1  1DAQ  
     1DAV  1DYM  1DYS  1E5J  1ECE  1EDG  1EG1  1EGZ  1F9D  1F9O  
     1FAE  1FBO  1FBW  1FCE  1G01  1G0C  1G87  1G9G  1G9J  1GA2  
     1GU3  1GZJ  1H0B  1H11  1H1N  1H2J  1H5V  1H8V  1HD5  1HF6  
     1IA6  1IA7  1IS9  1J83  1J84  1JS4  1K72  1KFG  1KS4  1KS5  
     1KS8  1KSC  1KSD  1KWF  1L1Y  1L2A  1L8F  1LF1  1NLR  1OA2  
     1OA3  1OA4  1OA7  1OA9  1OCQ  1OJI  1OJJ  1OJK  1OLQ  1OLR  
     1OVW  1QHZ  1QI0  1QI2  1TF4  1TML  1TVN  1TVP  1ULO  1ULP  
     1UT9  1UU4  1UU5  1UU6  1UWW  1V0A  1VJZ  1VRX  1W2U  1W3K  
     1W3L  1WC2  1WZZ  2A39  2A3H  2BOD  2BOE  2BOF  2BOG  2BV9  
     2BVD  2BW8  2BWA  2BWC  2CIP  2CIT  2CKR  2CKS  2DEP  2E0P  
     2E4T  2EEX  2EJ1  2ENG  2EO7  2EQD  2JEM  2JEN  2NLR  2OVW  
     2QNO  2UWA  2UWB  2UWC  2V38  2V3G  2VH9  2VI0  2WAB  2WAO  
     2ZUM  2ZUN  3A3H  3B7M  3ENG  3EZ8  3FW6  3GZK  3H2W  3H3K  
     3II1  3OVW  3TF4  4A3H  4ENG  4OVW  4TF4  5A3H  6A3H  7A3H  
     8A3H  
Reference
  Authors
  Title


  Journal
1  [PMID:14025219]
DATTA PK, HANSON KR, WHITAKER DR.
Improved procedures for preparation and characterization of
Myrothecium cellulase. 3. Molecular weight, amino acid composition,
terminal residues, and other properties.
Can. J. Biochem. Physiol. 41 (1963) 697-705.
Reference
  Authors
  Title
  Journal

2
Larner, J.
Other glucosidases.
In: Boyer, P.D., Lardy, H. and Myrback, K. (Eds.), The Enzymes, 2nd
ed., vol. 4, Academic Press, New York, 1960, p. 369-378.
Reference
  Authors
  Title

  Journal
  Organism
3  [PMID:13651124]
MYERS FL, NORTHCOTE DH.
Partial purification and some properties of a cellulase from Helix
pomatia.
Biochem. J. 71 (1959) 749-56.
Helix pomatia
Reference
  Authors
  Title

  Journal
4
Nishizawa, K. and Hashimoto, Y.
Cellulose splitting enzymes. VI. Difference in the specificities of
cellulase and beta-glucosidase from Irpex lacteus.
Arch. Biochem. Biophys. 81 (1959) 211-222.
Reference
  Authors
  Title

  Journal
5  [PMID:14000266]
WHITAKER DR, HANSON KR, DATTA PK.
Improved procedures for preparation and characterization of
myrothecium cellulase. 2. Purification procedures.
Can. J. Biochem. Physiol. 41 (1963) 671-96.
Reference
  Authors
  Title

  Journal
6
Hatfield, R. and Nevins, D.J.
Purification and properties of an endoglucanase isolated from the
cell walls of Zea mays seedlings.
Carbohydr. Res. 148 (1986) 265-278.
Reference
  Authors
  Title

  Journal
  Organism
7  [PMID:16665203]
Hatfield RD, Nevins DJ.
Hydrolytic Activity and Substrate Specificity of an Endoglucanase
from Zea mays Seedling Cell Walls.
Plant. Physiol. 83 (1987) 203-207.
Zea mays [GN:zma]
Reference
  Authors
  Title

  Journal
8
Inohue, M., Hayashgi, K. and Nevins, D.J.
Polypeptide characteristics and immunological properties of exo- and
endoglucanases purified from maize coleoptile cell walls.
J. Plant Physiol. 154 (1999) 334-340.
Other DBs ExplorEnz - The Enzyme Database: 3.2.1.4
IUBMB Enzyme Nomenclature: 3.2.1.4
ExPASy - ENZYME nomenclature database: 3.2.1.4
BRENDA, the Enzyme Database: 3.2.1.4
CAS: 9012-54-8

DBGET integrated database retrieval system