#=GF ID DHHC
#=GF AC PF01529.23
#=GF DE DHHC palmitoyltransferase
#=GF PI zf-DHHC;
#=GF AU Bateman A;0000-0002-6982-4660
#=GF SE Pfam-B_945 (release 4.0)
#=GF GA 28.70 28.70;
#=GF TC 28.70 28.70;
#=GF NC 28.60 28.60;
#=GF BM hmmbuild HMM.ann SEED.ann
#=GF SM hmmsearch -Z 61295632 -E 1000 --cpu 4 HMM pfamseq
#=GF TP Family
#=GF WK DHHC_domain
#=GF RN [1]
#=GF RM 15603741
#=GF RT Identification of PSD-95 palmitoylating enzymes.
#=GF RA Fukata M, Fukata Y, Adesnik H, Nicoll RA, Bredt DS;
#=GF RL Neuron. 2004;44:987-996.
#=GF RN [2]
#=GF RM 23034182
#=GF RT Analysis of substrate specificity of human DHHC protein
#=GF RT acyltransferases using a yeast expression system.
#=GF RA Ohno Y, Kashio A, Ogata R, Ishitomi A, Yamazaki Y, Kihara A;
#=GF RL Mol Biol Cell. 2012;23:4543-4551.
#=GF RN [3]
#=GF RM 26224664
#=GF RT The canonical DHHC motif is not absolutely required for the
#=GF RT activity of the yeast S-acyltransferases Swf1 and Pfa4.
#=GF RA Gonzalez Montoro A, Chumpen Ramirez S, Valdez Taubas J;
#=GF RL J Biol Chem. 2015;290:22448-22459.
#=GF RN [4]
#=GF RM 10231582
#=GF RT The drosophila STAM gene homolog is in a tight gene cluster, and
#=GF RT its expression correlates to that of the adjacent gene ial.
#=GF RA Mesilaty-Gross S, Reich A, Motro B, Wides R;
#=GF RL Gene 1999;231:173-186.
#=GF RN [5]
#=GF RM 12021275
#=GF RT Aph2, a protein with a zf-DHHC motif, interacts with c-Abl and
#=GF RT has pro-apoptotic activity.
#=GF RA Li B, Cong F, Tan CP, Wang SX, Goff SP;
#=GF RL J Biol Chem. 2002;277:28870-28876.
#=GF RN [6]
#=GF RM 16000296
#=GF RT DHHC9 and GCP16 constitute a human protein fatty acyltransferase
#=GF RT with specificity for H- and N-Ras.
#=GF RA Swarthout JT, Lobo S, Farh L, Croke MR, Greentree WK, Deschenes
#=GF RA RJ, Linder ME;
#=GF RL J Biol Chem. 2005;280:31141-31148.
#=GF RN [7]
#=GF RM 12370247
#=GF RT The yeast DHHC cysteine-rich domain protein Akr1p is a palmitoyl
#=GF RT transferase.
#=GF RA Roth AF, Feng Y, Chen L, Davis NG;
#=GF RL J Cell Biol. 2002;159:23-28.
#=GF RN [8]
#=GF RM 10490616
#=GF RT Erf2, a Novel Gene Product That Affects the Localization and
#=GF RT Palmitoylation of Ras2 in Saccharomyces cerevisiae.
#=GF RA Bartels DJ, Mitchell DA, Dong X, Deschenes RJ;
#=GF RL Mol Cell Biol 1999;19:6775-6787.
#=GF RN [9]
#=GF RM 16100337
#=GF RT The TIP GROWTH DEFECTIVE1 S-acyl transferase regulates plant
#=GF RT cell growth in Arabidopsis.
#=GF RA Hemsley PA, Kemp AC, Grierson CS;
#=GF RL Plant Cell. 2005;17:2554-2563.
#=GF DR INTERPRO; IPR001594;
#=GF DR SO; 0100021; polypeptide_conserved_region;
#=GF CC This entry refers to the DHHC domain, found in DHHC proteins
#=GF CC which are palmitoyltransferases [1]. Palmitoylation or, more
#=GF CC specifically S-acylation, plays important roles in the
#=GF CC regulation of protein localization, stability, and activity. It
#=GF CC is a post-translational protein modification that involves the
#=GF CC attachment of palmitic acid to Cys residues through a thioester
#=GF CC linkage. Protein acyltransferases (PATs), also known as
#=GF CC palmitoyltransferases, catalyze this reaction by transferring
#=GF CC the palmitoyl group from palmitoyl-CoA to the thiol group of
#=GF CC Cys residues. They are characterized by the presence of a
#=GF CC 50-residue-long domain called the DHHC domain, which in most
#=GF CC but not all cases is also cysteine-rich and gets its name from
#=GF CC a highly conserved DHHC signature tetrapeptide
#=GF CC (Asp-His-His-Cys). The Cys residue within the DHHC domain forms
#=GF CC a stable acyl intermediate and transfers the acyl chain to the
#=GF CC Cys residues of a target protein [2][3]. Some proteins
#=GF CC containing a DHHC domain include Drosophila DNZ1 protein [4],
#=GF CC Mouse Abl-philin 2 (Aph2) protein [5], Mammalian ZDHHC9 [6],
#=GF CC Yeast ankyrin repeat-containing protein AKR1 [7], Yeast Erf2
#=GF CC protein [8], and Arabidopsis thaliana tip growth defective 1
#=GF CC [9].
#=GF SQ 25560
#=GS F9X752_ZYMTI/1-89 AC F9X752.1
#=GS A0A061GXB1_THECC/150-252 AC A0A061GXB1.1
#=GS A0A5N5PRE0_PANHP/60-190 AC A0A5N5PRE0.1
#=GS A0A2K5IUY1_COLAP/153-305 AC A0A2K5IUY1.1
#=GS A0A0C2X147_9AGAM/93-222 AC A0A0C2X147.1
#=GS A0A1E5V1Q9_9POAL/168-298 AC A0A1E5V1Q9.1
#=GS T1IY67_STRMM/64-210 AC T1IY67.1
#=GS R0J0X2_SETT2/126-255 AC R0J0X2.1
#=GS A0A453IDX2_AEGTS/201-330 AC A0A453IDX2.1
#=GS A0A662XSM1_9STRA/120-243 AC A0A662XSM1.1
#=GS A0A2Y9GID1_NEOSC/126-287 AC A0A2Y9GID1.1
#=GS G8YAA2_PICSO/74-206 AC G8YAA2.1
#=GS A0A6P8FVL8_CLUHA/172-321 AC A0A6P8FVL8.1
#=GS B4H6D3_DROPE/112-242 AC B4H6D3.1
#=GS A0A1D6EM62_MAIZE/174-314 AC A0A1D6EM62.1
#=GS A0A0E0FB17_9ORYZ/350-457 AC A0A0E0FB17.1
#=GS A0A671QKP4_9TELE/132-268 AC A0A671QKP4.1
#=GS A0A2H5MX14_CITUN/148-289 AC A0A2H5MX14.1
#=GS A0A086TKP2_9FUNG/231-372 AC A0A086TKP2.1
#=GS M4BKD4_HYAAE/1-139 AC M4BKD4.1
#=GS A0A2Y9P747_DELLE/129-285 AC A0A2Y9P747.1
#=GS A0A0K0DP72_ANGCA/94-239 AC A0A0K0DP72.1
#=GS G0QYL7_ICHMG/149-275 AC G0QYL7.1
#=GS A0A672V1H8_STRHB/99-224 AC A0A672V1H8.1
#=GS A0A6J2T599_DROLE/92-234 AC A0A6J2T599.1
#=GS F0ULX9_AJEC8/161-285 AC F0ULX9.1
#=GS A0A1D3RIS8_PLAMA/106-232 AC A0A1D3RIS8.1
#=GS A0A6P8YJX9_DROAB/124-251 AC A0A6P8YJX9.1
#=GS A0A6I9I772_VICPA/92-218 AC A0A6I9I772.1
#=GS A0A3L6QM92_PANMI/107-237 AC A0A3L6QM92.1
#=GS A0A2A2KDT7_9BILA/47-183 AC A0A2A2KDT7.1
#=GS A0A179UPY5_BLAGS/151-304 AC A0A179UPY5.1
#=GS A0A669PUY2_PHACC/134-263 AC A0A669PUY2.1
#=GS A0A087R672_APTFO/162-295 AC A0A087R672.1
#=GS A0A1R3JUS3_9ROSI/120-263 AC A0A1R3JUS3.1
#=GS A0A177C660_9PLEO/114-238 AC A0A177C660.1
#=GS A0A2U4CPM6_TURTR/160-289 AC A0A2U4CPM6.1
#=GS A0A0K0EWV8_STRVS/115-267 AC A0A0K0EWV8.1
#=GS L8GCE9_PSED2/122-251 AC L8GCE9.1
#=GS A0A6J0A3B4_ACIJB/92-242 AC A0A6J0A3B4.1
#=GS A0A0C3F6N8_PILCF/125-257 AC A0A0C3F6N8.1
#=GS A0A0J8QQT9_COCIT/150-275 AC A0A0J8QQT9.1
#=GS A0A383Z6E2_BALAS/136-265 AC A0A383Z6E2.1
#=GS ZDH20_BOVIN/121-247 AC Q0VC89.1
#=GS B7PS34_IXOSC/124-249 AC B7PS34.1
#=GS A0A5K4E974_SCHMA/18-186 AC A0A5K4E974.1
#=GS K1Y8J5_MARBU/421-551 AC K1Y8J5.1
#=GS A0A5D2VKF6_GOSMU/142-273 AC A0A5D2VKF6.1
#=GS A0A0S3SN92_PHAAN/137-267 AC A0A0S3SN92.1
#=GS A0A3P9CA55_9CICH/148-272 AC A0A3P9CA55.1
#=GS A0A7E6ERA7_OCTVU/125-278 AC A0A7E6ERA7.1
#=GS A0A4Q4UAN4_9PEZI/152-302 AC A0A4Q4UAN4.1
#=GS A0A565C744_9BRAS/142-273 AC A0A565C744.1
#=GS A0A158NH53_ATTCE/134-302 AC A0A158NH53.1
#=GS B1N5J3_ENTHI/141-273 AC B1N5J3.1
#=GS A0A4C2A4C1_EUMVA/131-262 AC A0A4C2A4C1.1
#=GS M2U4B8_COCH5/153-316 AC M2U4B8.1
#=GS A0A6I9LSQ8_PERMB/99-224 AC A0A6I9LSQ8.1
#=GS A0A091VNB8_NIPNI/150-289 AC A0A091VNB8.1
#=GS A0A137PGK5_CONC2/54-188 AC A0A137PGK5.1
#=GS A0A4W5PTF5_9TELE/122-277 AC A0A4W5PTF5.1
#=GS A0A3Q3FVE0_9LABR/99-224 AC A0A3Q3FVE0.1
#=GS A0A5E4EAK5_PRUDU/155-306 AC A0A5E4EAK5.1
#=GS A0A103YKR9_CYNCS/377-510 AC A0A103YKR9.1
#=GS A0A3Q4M5N5_NEOBR/99-224 AC A0A3Q4M5N5.1
#=GS G0QPP3_ICHMG/85-222 AC G0QPP3.1
#=GS A0A1U7SUD8_CARSF/85-218 AC A0A1U7SUD8.1
#=GS A0A074W618_9PEZI/177-301 AC A0A074W618.1
#=GS A0A367L6C3_9HYPO/151-299 AC A0A367L6C3.1
#=GS A0A1J9P7F3_9EURO/151-304 AC A0A1J9P7F3.1
#=GS A0A2K5HQ36_COLAP/99-224 AC A0A2K5HQ36.1
#=GS F7D678_MONDO/164-293 AC F7D678.2
#=GS M2NFX2_BAUPA/158-287 AC M2NFX2.1
#=GS A0A5E4NLF0_9HEMI/94-219 AC A0A5E4NLF0.1
#=GS A0A397Z3N8_BRACM/144-275 AC A0A397Z3N8.1
#=GS A0A267EQB0_9PLAT/197-323 AC A0A267EQB0.1
#=GS A0A427YNJ7_9TREE/143-271 AC A0A427YNJ7.1
#=GS A0A5E4F031_PRUDU/151-292 AC A0A5E4F031.1
#=GS A0A316YQP0_9BASI/1068-1211 AC A0A316YQP0.1
#=GS A0A319C4A2_9EURO/402-531 AC A0A319C4A2.1
#=GS A0A384BAT8_BALAS/99-224 AC A0A384BAT8.1
#=GS A0A3Q7FQ35_SOLLC/149-280 AC A0A3Q7FQ35.1
#=GS A0A485PB36_LYNPA/116-260 AC A0A485PB36.1
#=GS A0A1D2MRG9_ORCCI/157-285 AC A0A1D2MRG9.1
#=GS A0A091L4M3_CATAU/85-218 AC A0A091L4M3.1
#=GS A0A078B4X9_STYLE/85-216 AC A0A078B4X9.1
#=GS A0A446QRT9_TRITD/146-261 AC A0A446QRT9.1
#=GS G3ARQ4_SPAPN/53-181 AC G3ARQ4.1
#=GS A0A2Y9M3J1_DELLE/417-553 AC A0A2Y9M3J1.1
#=GS A0A2P6P7Y1_ROSCH/126-255 AC A0A2P6P7Y1.1
#=GS ZD12B_DANRE/97-223 AC E7F021.1
#=GS A0A6J0KXR5_RAPSA/355-488 AC A0A6J0KXR5.1
#=GS A0A4V6A947_POPAL/134-264 AC A0A4V6A947.1
#=GS A0A1E3P5T6_WICAA/133-264 AC A0A1E3P5T6.1
#=GS A0A210PHG9_MIZYE/117-243 AC A0A210PHG9.1
#=GS A0A2A3EKM2_APICC/408-547 AC A0A2A3EKM2.1
#=GS A0A4U6UBD0_SETVI/100-239 AC A0A4U6UBD0.1
#=GS A0A445J6R8_GLYSO/120-249 AC A0A445J6R8.1
#=GS A0A452G0R6_CAPHI/468-579 AC A0A452G0R6.1
#=GS A0A6I8WDS6_DROPS/94-219 AC A0A6I8WDS6.1
#=GS A0A670JJL8_PODMU/90-234 AC A0A670JJL8.1
#=GS A0A067E7P0_CITSI/91-124 AC A0A067E7P0.1
#=GS A0A6P7M386_BETSP/118-273 AC A0A6P7M386.1
#=GS A0A6P8XV84_DROAB/257-399 AC A0A6P8XV84.1
#=GS A0A6I9IJP9_VICPA/99-224 AC A0A6I9IJP9.1
#=GS A0A5J4Z3B7_PORPP/131-271 AC A0A5J4Z3B7.1
#=GS A0A1B7MVD6_9AGAM/376-510 AC A0A1B7MVD6.1
#=GS A0A3B0J350_DROGU/124-254 AC A0A3B0J350.1
#=GS A0A6P4E6N6_DRORH/124-263 AC A0A6P4E6N6.1
#=GS G0S146_CHATD/364-494 AC G0S146.1
#=GS A0A1D3U8C3_9APIC/207-334 AC A0A1D3U8C3.1
#=GS A0A1U7S0U2_ALLSI/162-291 AC A0A1U7S0U2.1
#=GS A0A2I0B140_9ASPA/72-201 AC A0A2I0B140.1
#=GS A0A2R6QBY7_ACTCC/154-311 AC A0A2R6QBY7.1
#=GS R7T6R7_CAPTE/142-270 AC R7T6R7.1
#=GS A0A6J2M274_9CHIR/99-224 AC A0A6J2M274.1
#=GS C0NMD1_AJECG/393-531 AC C0NMD1.1
#=GS F6PJF9_MACMU/193-322 AC F6PJF9.3
#=GS A0A482WLS1_LAOST/113-255 AC A0A482WLS1.1
#=GS A0A6P6FXS1_ZIZJJ/143-240 AC A0A6P6FXS1.1
#=GS A0A6P9A4I7_THRPL/125-250 AC A0A6P9A4I7.1
#=GS H3CLW6_TETNG/137-267 AC H3CLW6.1
#=GS A0A2R6XRY9_MARPO/157-308 AC A0A2R6XRY9.1
#=GS A0A0K9QCJ4_SPIOL/74-225 AC A0A0K9QCJ4.1
#=GS A0A1V6S2G5_9EURO/150-296 AC A0A1V6S2G5.1
#=GS A0A6P6EIM9_OCTDE/384-520 AC A0A6P6EIM9.1
#=GS A0A6P5B793_BOSIN/150-289 AC A0A6P5B793.1
#=GS A0A6Q2Z3Y2_ESOLU/265-401 AC A0A6Q2Z3Y2.1
#=GS A0A6J0UZK3_9SAUR/59-188 AC A0A6J0UZK3.1
#=GS A0A445E2I4_ARAHY/102-242 AC A0A445E2I4.1
#=GS A0A6I8TDK2_AEDAE/418-558 AC A0A6I8TDK2.1
#=GS A0A0K9NN50_ZOSMR/373-506 AC A0A0K9NN50.1
#=GS A0A1R3HYK3_COCAP/358-490 AC A0A1R3HYK3.1
#=GS A0A2J8XXU0_PONAB/151-305 AC A0A2J8XXU0.1
#=GS A0A6J3S141_TURTR/126-254 AC A0A6J3S141.1
#=GS A0A3B0J517_DROGU/425-561 AC A0A3B0J517.1
#=GS A0A067JY22_JATCU/120-249 AC A0A067JY22.1
#=GS A0A3Q1BHE7_AMPOC/339-472 AC A0A3Q1BHE7.1
#=GS A0A540MLZ4_MALBA/108-230 AC A0A540MLZ4.1
#=GS A0A2K5S9I6_CEBIM/159-288 AC A0A2K5S9I6.1
#=GS A0A4W4FL77_ELEEL/98-236 AC A0A4W4FL77.1
#=GS J9IA99_9SPIT/82-222 AC J9IA99.1
#=GS A0A3S5WGI8_9MAGN/138-267 AC A0A3S5WGI8.1
#=GS A0A2K6UGU6_SAIBB/99-224 AC A0A2K6UGU6.1
#=GS A0A4Z1L551_9HELO/444-574 AC A0A4Z1L551.1
#=GS A0A6P8XDE5_DROAB/82-215 AC A0A6P8XDE5.1
#=GS I1JCY3_SOYBN/145-276 AC I1JCY3.1
#=GS A0A1R2BY23_9CILI/141-268 AC A0A1R2BY23.1
#=GS A0A137P958_CONC2/220-355 AC A0A137P958.1
#=GS A0A4U5P9T0_STECR/119-264 AC A0A4U5P9T0.1
#=GS A0A6G0WM55_9STRA/367-482 AC A0A6G0WM55.1
#=GS L9JBL0_TUPCH/110-298 AC L9JBL0.1
#=GS J3LIF2_ORYBR/90-216 AC J3LIF2.1
#=GS A0A4S8L3R2_DENBC/111-248 AC A0A4S8L3R2.1
#=GS Q22BC3_TETTS/349-485 AC Q22BC3.3
#=GS A0A0D8XWI6_DICVI/117-218 AC A0A0D8XWI6.1
#=GS A0A2P6SJB5_ROSCH/91-218 AC A0A2P6SJB5.1
#=GS A0A6G0T3S5_APHGL/125-288 AC A0A6G0T3S5.1
#=GS A0A086SVQ4_ACRC1/118-268 AC A0A086SVQ4.1
#=GS A0A6J2HUV0_9PASS/137-262 AC A0A6J2HUV0.1
#=GS A0A446YJF5_TRITD/128-257 AC A0A446YJF5.1
#=GS A0A6J0WSR7_ODOVR/150-305 AC A0A6J0WSR7.1
#=GS A0A1J7J075_9PEZI/540-677 AC A0A1J7J075.1
#=GS A0A673K4H0_9TELE/93-229 AC A0A673K4H0.1
#=GS A0A6J2N320_9CHIR/99-224 AC A0A6J2N320.1
#=GS A0A6P6MX10_CARAU/444-580 AC A0A6P6MX10.1
#=GS A0A078AWI7_STYLE/1-123 AC A0A078AWI7.1
#=GS A0A3Q7GZH2_SOLLC/149-280 AC A0A3Q7GZH2.1
#=GS A0A482XR05_LAOST/160-297 AC A0A482XR05.1
#=GS A0A1A0HCI7_9ASCO/166-302 AC A0A1A0HCI7.1
#=GS A0A0K0DYK5_STRER/124-265 AC A0A0K0DYK5.1
#=GS A0A1S3RHA3_SALSA/226-265 AC A0A1S3RHA3.1
#=GS A0A674P685_TAKRU/241-375 AC A0A674P685.1
#=GS A0A072UR39_MEDTR/199-331 AC A0A072UR39.1
#=GS A0A671TNC7_SPAAU/98-156 AC A0A671TNC7.1
#=GS A0A4W4HTN6_ELEEL/115-244 AC A0A4W4HTN6.1
#=GS A0A402FR05_9SAUR/119-244 AC A0A402FR05.1
#=GS A0A3N7GQN0_POPTR/145-186 AC A0A3N7GQN0.1
#=GS A0A6P3XK52_DINQU/413-552 AC A0A6P3XK52.1
#=GS A0A3P9BPA2_9CICH/424-560 AC A0A3P9BPA2.1
#=GS A0A6J3GDF3_SAPAP/121-247 AC A0A6J3GDF3.1
#=GS A0A6J2L028_9CHIR/253-384 AC A0A6J2L028.1
#=GS W9QV74_9ROSA/160-291 AC W9QV74.1
#=GS A0A017SHY5_9EURO/125-254 AC A0A017SHY5.1
#=GS A0A161V3P3_9PEZI/115-244 AC A0A161V3P3.1
#=GS A0A3R7M6T4_TRYRA/91-227 AC A0A3R7M6T4.1
#=GS A0A2I4AUI6_9TELE/99-224 AC A0A2I4AUI6.1
#=GS A0A670YFA6_PSETE/394-530 AC A0A670YFA6.1
#=GS A0A5E4NJ28_9HEMI/66-208 AC A0A5E4NJ28.1
#=GS A0A7E6DRA6_9CHIR/183-312 AC A0A7E6DRA6.1
#=GS V5DBY9_TRYCR/1-171 AC V5DBY9.1
#=GS A0A1W4XIZ8_AGRPL/84-226 AC A0A1W4XIZ8.1
#=GS A0A1S4DRH6_TOBAC/373-506 AC A0A1S4DRH6.1
#=GS A0A061DC06_BABBI/105-243 AC A0A061DC06.1
#=GS A0A484GNI1_SOUCH/254-388 AC A0A484GNI1.1
#=GS A0A5D2WFS3_GOSMU/173-313 AC A0A5D2WFS3.1
#=GS A0A395NP04_TRIAR/80-213 AC A0A395NP04.1
#=GS A0A341AEE4_NEOAA/115-271 AC A0A341AEE4.1
#=GS A0A653H0W9_9APIC/93-245 AC A0A653H0W9.1
#=GS A0A3Q3B665_KRYMA/98-242 AC A0A3Q3B665.1
#=GS A0A674HDS8_TAEGU/581-718 AC A0A674HDS8.1
#=GS A0DRQ8_PARTE/325-456 AC A0DRQ8.1
#=GS A0A671DHM0_RHIFE/116-240 AC A0A671DHM0.1
#=GS A0A2K5SDC8_CEBIM/187-316 AC A0A2K5SDC8.1
#=GS A0A1A8Z4Y6_9APIC/21-157 AC A0A1A8Z4Y6.1
#=GS M4CJJ6_BRARP/156-296 AC M4CJJ6.1
#=GS A0A182Y9U2_ANOST/1802-1944 AC A0A182Y9U2.1
#=GS F2EJY1_HORVV/146-276 AC F2EJY1.1
#=GS A0A0K3C963_RHOTO/901-1091 AC A0A0K3C963.1
#=GS A0A1D6HFL4_MAIZE/146-197 AC A0A1D6HFL4.1
#=GS A0A445F649_GLYSO/137-267 AC A0A445F649.1
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