+C Secretion system KO
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Secretion System - Pseudomonas aeruginosa DK2
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AType I secretion system
B alpha-Hemolysin/cyclolysin transporter
C PADK2_26360 putative outer membrane protein precursor K12340 tolC; outer membrane protein
C PADK2_24390 outer membrane protein precursor K12340 tolC; outer membrane protein
B RTX toxin transporter
C PADK2_26360 putative outer membrane protein precursor K12340 tolC; outer membrane protein
C PADK2_24390 outer membrane protein precursor K12340 tolC; outer membrane protein
B Multiple protein transporter
C PADK2_07235 transport protein HasD K12536 hasD; ATP-binding cassette, subfamily C, type I secretion system permease/ATPase
C PADK2_19420 alkaline protease secretion protein AprD K12536 hasD; ATP-binding cassette, subfamily C, type I secretion system permease/ATPase
C PADK2_07240 metalloprotease secretion protein K12537 hasE; membrane fusion protein, type I secretion system
C PADK2_19415 alkaline protease secretion protein AprE K12537 hasE; membrane fusion protein, type I secretion system
C PADK2_07245 outer membrane protein HasF K12538 hasF; outer membrane protein, type I secretion system
C PADK2_19410 alkaline protease secretion outer membrane protein AprF precursor K12538 hasF; outer membrane protein, type I secretion system
C PADK2_26360 putative outer membrane protein precursor K12340 tolC; outer membrane protein
C PADK2_24390 outer membrane protein precursor K12340 tolC; outer membrane protein
B Adhesin protein transporter
C PADK2_16270 ATP-binding/permease fusion ABC transporter K12541 lapB; ATP-binding cassette, subfamily C, bacterial LapB
C PADK2_16265 putative secretion protein K12542 lapC; membrane fusion protein, adhesin transport system
C PADK2_16275 putative outer membrane protein precursor K12543 lapE; outer membrane protein, adhesin transport system
B RaxAB-RaxC type I secretion system protein
C PADK2_26360 putative outer membrane protein precursor K12340 tolC; outer membrane protein
C PADK2_24390 outer membrane protein precursor K12340 tolC; outer membrane protein
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AType II secretion system
B General secretion pathway protein
C PADK2_08815 secretion protein XcpP K02452 gspC; general secretion pathway protein C
C PADK2_16315 XphA K02452 gspC; general secretion pathway protein C
C PADK2_22255 hypothetical protein K02452 gspC; general secretion pathway protein C
C PADK2_08810 general secretion pathway protein D K02453 gspD; general secretion pathway protein D
C PADK2_16310 secretion protein XqhA K02453 gspD; general secretion pathway protein D
C PADK2_22225 general secretion pathway protein D K02453 gspD; general secretion pathway protein D
C PADK2_08820 general secretion pathway protein E K02454 gspE; general secretion pathway protein E [EC:7.4.2.8]
C PADK2_22220 putative type II secretion system protein K02454 gspE; general secretion pathway protein E [EC:7.4.2.8]
C PADK2_27570 putative secretion pathway ATPase K02454 gspE; general secretion pathway protein E [EC:7.4.2.8]
C PADK2_23810 hypothetical protein K02454 gspE; general secretion pathway protein E [EC:7.4.2.8]
C PADK2_11270 putative type II secretion system protein K02455 gspF; general secretion pathway protein F
C PADK2_08825 general secretion pathway protein F K02455 gspF; general secretion pathway protein F
C PADK2_22215 putative type II secretion system protein K02455 gspF; general secretion pathway protein F
C PADK2_11275 general secretion pathway protein G K02456 gspG; general secretion pathway protein G
C PADK2_11280 putative type II secretion system protein K02456 gspG; general secretion pathway protein G
C PADK2_08830 general secretion pathway protein G K02456 gspG; general secretion pathway protein G
C PADK2_22245 HxcT pseudopilin K02456 gspG; general secretion pathway protein G
C PADK2_08835 general secretion pathway protein H K02457 gspH; general secretion pathway protein H
C PADK2_22260 HxcU putative pseudopilin K02457 gspH; general secretion pathway protein H
C PADK2_11285 type II secretion system protein K02458 gspI; general secretion pathway protein I
C PADK2_08840 general secretion pathway protein I K02458 gspI; general secretion pathway protein I
C PADK2_22250 general secretion pathway protein I K02458 gspI; general secretion pathway protein I
C PADK2_11290 putative type II secretion system protein K02459 gspJ; general secretion pathway protein J
C PADK2_08845 general secretion pathway protein J K02459 gspJ; general secretion pathway protein J
C PADK2_22265 HxcW putative pseudopilin K02459 gspJ; general secretion pathway protein J
C PADK2_08850 general secretion pathway protein K K02460 gspK; general secretion pathway protein K
C PADK2_22240 putative type II secretion system protein K02460 gspK; general secretion pathway protein K
C PADK2_08855 general secretion pathway protein L K02461 gspL; general secretion pathway protein L
C PADK2_22235 type II secretion system protein K02461 gspL; general secretion pathway protein L
C PADK2_08860 general secretion pathway protein M K02462 gspM; general secretion pathway protein M
C PADK2_22230 type II secretion system protein K02462 gspM; general secretion pathway protein M
C PADK2_23590 type 4 prepilin peptidase PilD K02654 pilD; leader peptidase (prepilin peptidase) / N-methyltransferase [EC:3.4.23.43 2.1.1.-]
B Pilin secretion/fimbrial assembly protein
C PADK2_23560 type IV pilin structural subunit K02650 pilA; type IV pilus assembly protein PilA
C PADK2_23565 type 4 fimbrial biogenesis protein PilB K02652 pilB; type IV pilus assembly protein PilB
C PADK2_23585 pilin biogenesis protein K02653 pilC; type IV pilus assembly protein PilC
C PADK2_23590 type 4 prepilin peptidase PilD K02654 pilD; leader peptidase (prepilin peptidase) / N-methyltransferase [EC:3.4.23.43 2.1.1.-]
C PADK2_24210 type 4 fimbrial biogenesis protein PilE K02655 pilE; type IV pilus assembly protein PilE
C PADK2_05300 type IV pilus biogenesis/stability protein PilW K02656 pilF; type IV pilus assembly protein PilF
C PADK2_02055 twitching motility protein PilG K02657 pilG; twitching motility two-component system response regulator PilG
C PADK2_02060 twitching motility protein PilH K02658 pilH; twitching motility two-component system response regulator PilH
C PADK2_02065 twitching motility protein PilI K02659 pilI; twitching motility protein PilI
C PADK2_02070 twitching motility protein PilJ K02660 pilJ; twitching motility protein PilJ
C PADK2_02075 methyltransferase PilK K02661 pilK; type IV pilus assembly protein PilK
C PADK2_26715 type IV pilus assembly protein PilM K02662 pilM; type IV pilus assembly protein PilM
C PADK2_26710 type 4 fimbrial biogenesis protein PilN K02663 pilN; type IV pilus assembly protein PilN
C PADK2_26705 type 4 fimbrial biogenesis protein PilO K02664 pilO; type IV pilus assembly protein PilO
C PADK2_26700 type 4 fimbrial biogenesis protein PilP K02665 pilP; type IV pilus assembly protein PilP
C PADK2_26695 type 4 fimbrial biogenesis outer membrane protein PilQ precursor K02666 pilQ; type IV pilus assembly protein PilQ
C PADK2_01990 twitching motility protein PilT K02669 pilT; twitching motility protein PilT
C PADK2_01995 twitching motility protein PilU K02670 pilU; twitching motility protein PilU
C PADK2_24185 type IV pilus modification protein PilV K02671 pilV; type IV pilus assembly protein PilV
C PADK2_24190 type 4 fimbrial biogenesis protein PilW K02672 pilW; type IV pilus assembly protein PilW
C PADK2_24195 type 4 fimbrial biogenesis protein PilX K02673 pilX; type IV pilus assembly protein PilX
C PADK2_24200 type 4 fimbrial biogenesis protein PilY1 K02674 pilY1; type IV pilus assembly protein PilY1
C PADK2_09555 type 4 fimbrial biogenesis protein PilZ K02676 pilZ; type IV pilus assembly protein PilZ
C PADK2_16555 pilin biosynthetic protein K27041 fimL; scaffold protein FimL
C PADK2_24175 type 4 fimbrial biogenesis protein FimT K08084 fimT; type IV fimbrial biogenesis protein FimT
C PADK2_24180 type 4 fimbrial biogenesis protein FimU K08085 fimU; type IV fimbrial biogenesis protein FimU
C PADK2_08760 Motility protein FimV K08086 fimV; pilus assembly protein FimV
B Tight adherence (Tad) export apparatus
C PADK2_22380 Flp prepilin peptidase A, FppA K02278 cpaA; prepilin peptidase CpaA [EC:3.4.23.43]
C PADK2_22430 Flp pilus assembly protein CpaB K02279 cpaB; pilus assembly protein CpaB
C PADK2_22425 putative type II secretion system protein K02280 cpaC; pilus assembly protein CpaC
C PADK2_22420 TadZ K02282 cpaE; pilus assembly protein CpaE
C PADK2_22415 TadA ATPase K02283 cpaF; pilus assembly protein CpaF [EC:7.4.2.8]
C PADK2_22410 hypothetical protein K12510 tadB; tight adherence protein B
C PADK2_22405 TadC K12511 tadC; tight adherence protein C
C PADK2_22400 TadD K27079 tadD; tight adherence protein D
C PADK2_22390 TadG K27080 tadG; tight adherence protein G
C PADK2_22435 type IVb pilin, Flp K02651 flp; pilus assembly protein Flp/PilA
B Toxin co-regulated pilus biosynthesis protein
B MSHA pilus biogenesis protein
B Type IVa pilus homolog protein
C PADK2_23590 type 4 prepilin peptidase PilD K02654 pilD; leader peptidase (prepilin peptidase) / N-methyltransferase [EC:3.4.23.43 2.1.1.-]
C PADK2_01990 twitching motility protein PilT K02669 pilT; twitching motility protein PilT
C PADK2_01975 YGGT family protein K02221 yggT; YggT family protein
B Archaeal flagellar protein
B Competence-related DNA transformation transporter (DNA-T) core components
C PADK2_08630 putative competence protein K02237 comEA; competence protein ComEA
C PADK2_09435 DNA internalization-related competence protein ComEC/Rec2 K02238 comEC; competence protein ComEC
B Competence-related DNA transformation transporter (DNA-T) related proteins
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AType III secretion system
B Type III secretion core apparatus
C PADK2_17115 Type III secretion outer membrane protein PscC precursor K03219 yscC; type III secretion protein C
C PADK2_17110 type III export protein PscD K03220 yscD; type III secretion protein D
C PADK2_17100 type III export protein PscF K03221 yscF; type III secretion protein F
C PADK2_17085 type III export protein PscI K04053 yscI; type III secretion protein I
C PADK2_17080 pscJ type III export protein K03222 yscJ; type III secretion protein J
C PADK2_17075 type III export protein PscK K04054 yscK; type III secretion protein K
C PADK2_17070 type III secretion system protein K03223 yscL; type III secretion protein L
C PADK2_17215 type III secretion system ATPase K03224 yscN; ATP synthase in type III secretion protein N [EC:7.4.2.8]
C PADK2_17220 translocation protein in type III secretion K04056 yscO; type III secretion protein O
C PADK2_17225 translocation protein in type III secretion K04057 yscP; type III secretion protein P
C PADK2_17230 type III secretion system protein K03225 yscQ; type III secretion protein Q
C PADK2_17235 type III secretion system protein YscR K03226 yscR; type III secretion protein R
C PADK2_17240 putative translocation protein in type III secretion K03227 yscS; type III secretion protein S
C PADK2_17245 translocation protein in type III secretion K03228 yscT; type III secretion protein T
C PADK2_17250 translocation protein in type III secretion K03229 yscU; type III secretion protein U
C PADK2_17185 type III secretory apparatus protein PcrD K03230 yscV; type III secretion protein V
C PADK2_17210 Type III secretion outer membrane protein PopN precursor K04058 yscW; type III secretion protein W
C PADK2_17195 Yop proteins translocation protein X K04059 yscX; type III secretion protein X
B Type III secretion related proteins
C PADK2_17120 type III export apparatus protein K04049 yscB; type III secretion protein B
C PADK2_17105 type III export protein PscE K04050 yscE; type III secretion protein E
C PADK2_17095 type III secretion protein, YscG family K04051 yscG; type III secretion protein G
C PADK2_17090 type III export protein PscH K04052 yscH; type III secretion protein H
C PADK2_17190 hypothetical protein K04060 yscY; type III secretion protein Y
B Flagellar export apparatus
C PADK2_18465 flhA; flagellar biosynthesis protein FlhA K02400 flhA; flagellar biosynthesis protein FlhA
C PADK2_18480 flhB; flagellar biosynthesis protein FlhB K02401 flhB; flagellar biosynthesis protein FlhB
C PADK2_18360 putative FlhB domain protein K04061 flhB2; flagellar biosynthesis protein
C PADK2_20175 fliF; flagellar MS-ring protein K02409 fliF; flagellar M-ring protein FliF
C PADK2_20165 fliH; flagellar assembly protein H K02411 fliH; flagellar assembly protein FliH
C PADK2_20160 fliI; flagellum-specific ATP synthase K02412 fliI; flagellum-specific ATP synthase [EC:7.4.2.8]
C PADK2_18505 fliN; flagellar motor switch protein K02417 fliN; flagellar motor switch protein FliN
C PADK2_18500 flagellar protein FliO K02418 fliO; flagellar protein FliO/FliZ
C PADK2_18495 fliP; flagellar biosynthesis protein FliP K02419 fliP; flagellar biosynthesis protein FliP
C PADK2_18490 fliQ; flagellar biosynthesis protein FliQ K02420 fliQ; flagellar biosynthesis protein FliQ
C PADK2_18485 fliR; flagellar biosynthesis protein FliR K02421 fliR; flagellar biosynthesis protein FliR
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AType IV secretion system
B Conjugal DNA-protein transfer (VirB/D) protein
B DNA uptake competence system protein
B Conjugal transfer pilus assembly protein
C PADK2_12290 Cobyrinic acid a,c-diamide synthase K12055 K12055; chromosome partitioning related protein ParA
C PADK2_23660 chromosome partitioning related protein K12055 K12055; chromosome partitioning related protein ParA
C PADK2_11915 hypothetical protein K12070 traI; conjugal transfer pilus assembly protein TraI
C PADK2_24125 relaxase K12070 traI; conjugal transfer pilus assembly protein TraI
C PADK2_23970 TraG/TraD family protein K12071 traD; conjugal transfer pilus assembly protein TraD
C PADK2_12060 hypothetical protein K12071 traD; conjugal transfer pilus assembly protein TraD
C PADK2_05650 thiol:disulfide interchange protein DsbC K03981 dsbC; thiol:disulfide interchange protein DsbC [EC:5.3.4.1]
B Pertussis toxin exporter
B Cag pathogenicity island (PAI) protein
B Icm/Dot secretion system protein
B Trb secretion system protein
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AType V secretion system
B Autotransporter-1 (AT-1) family
C PADK2_06690 serine protease K12685 ssp; subtilase-type serine protease [EC:3.4.21.-]
C PADK2_27055 esterase EstA K12686 apeE; outer membrane lipase/esterase
B Autotransporter-2 (AT-2) family
B Two-partner secretion (TPS) family
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AType VI secretion system
B Imp/Vas secretion system core components
C PADK2_00405 hypothetical protein K11891 impL; type VI secretion system protein ImpL
C PADK2_13465 type VI secretion system protein ImpL K11891 impL; type VI secretion system protein ImpL
C PADK2_17360 type VI secretion protein IcmF K11891 impL; type VI secretion system protein ImpL
C PADK2_00410 hypothetical protein K11892 impK; type VI secretion system protein ImpK
C PADK2_13460 type VI secretion system protein ImpK K11892 impK; type VI secretion system protein ImpK
C PADK2_17365 type VI secretion system protein ImpK K11892 impK; type VI secretion system protein ImpK
C PADK2_00415 hypothetical protein K11893 impJ; type VI secretion system protein ImpJ
C PADK2_13455 hypothetical protein K11893 impJ; type VI secretion system protein ImpJ
C PADK2_17370 hypothetical protein K11893 impJ; type VI secretion system protein ImpJ
C PADK2_00465 hypothetical protein K11895 impH; type VI secretion system protein ImpH
C PADK2_13420 hypothetical protein K11895 impH; type VI secretion system protein ImpH
C PADK2_17400 hypothetical protein K11895 impH; type VI secretion system protein ImpH
C PADK2_00460 type VI secretion system protein ImpG K11896 impG; type VI secretion system protein ImpG
C PADK2_13425 type VI secretion system protein ImpG K11896 impG; type VI secretion system protein ImpG
C PADK2_17405 hypothetical protein K11896 impG; type VI secretion system protein ImpG
C PADK2_00455 hypothetical protein K11897 impF; type VI secretion system protein ImpF
C PADK2_00440 hypothetical protein K11900 impC; type VI secretion system protein ImpC
C PADK2_13440 hypothetical protein K11900 impC; type VI secretion system protein ImpC
C PADK2_17415 hypothetical protein K11900 impC; type VI secretion system protein ImpC
C PADK2_00435 hypothetical protein K11901 impB; type VI secretion system protein ImpB
C PADK2_13445 hypothetical protein K11901 impB; type VI secretion system protein ImpB
C PADK2_17420 hypothetical protein K11901 impB; type VI secretion system protein ImpB
C PADK2_00430 hypothetical protein K11902 impA; type VI secretion system protein ImpA
C PADK2_13470 hypothetical protein K11902 impA; type VI secretion system protein ImpA
C PADK2_00445 hypothetical protein K11903 hcp; type VI secretion system secreted protein Hcp
C PADK2_13435 hypothetical protein K11903 hcp; type VI secretion system secreted protein Hcp
C PADK2_01310 secreted protein Hcp K11903 hcp; type VI secretion system secreted protein Hcp
C PADK2_18160 secreted protein Hcp K11903 hcp; type VI secretion system secreted protein Hcp
C PADK2_27865 secreted protein Hcp K11903 hcp; type VI secretion system secreted protein Hcp
C PADK2_26945 hypothetical protein K11904 vgrG; type VI secretion system secreted protein VgrG
C PADK2_01305 hypothetical protein K11904 vgrG; type VI secretion system secreted protein VgrG
C PADK2_07815 hypothetical protein K11904 vgrG; type VI secretion system secreted protein VgrG
C PADK2_18165 hypothetical protein K11904 vgrG; type VI secretion system secreted protein VgrG
C PADK2_27860 hypothetical protein K11904 vgrG; type VI secretion system secreted protein VgrG
C PADK2_11220 hypothetical protein K11904 vgrG; type VI secretion system secreted protein VgrG
C PADK2_00495 hypothetical protein K11904 vgrG; type VI secretion system secreted protein VgrG
C PADK2_00475 hypothetical protein K11904 vgrG; type VI secretion system secreted protein VgrG
C PADK2_13400 hypothetical protein K11904 vgrG; type VI secretion system secreted protein VgrG
C PADK2_17410 hypothetical protein K11905 K11905; type VI secretion system protein
C PADK2_00420 putative lipoprotein K11906 vasD; type VI secretion system protein VasD
C PADK2_17375 putative lipoprotein K11906 vasD; type VI secretion system protein VasD
C PADK2_00470 putative ClpA/B-type chaperone K11907 vasG; type VI secretion system protein VasG
C PADK2_13415 ClpA/B-type protease K11907 vasG; type VI secretion system protein VasG
C PADK2_17395 putative ClpA/B-type protease K11907 vasG; type VI secretion system protein VasG
C PADK2_17425 hypothetical protein K11910 vasJ; type VI secretion system protein VasJ
B Imp/Vas secretion system related proteins
C PADK2_00400 type VI secretion system protein ImpM K11890 impM; type VI secretion system protein ImpM
C PADK2_17380 hypothetical protein K11894 impI; type VI secretion system protein ImpI
C PADK2_00450 putative secretion protein K11898 impE; type VI secretion system protein ImpE
C PADK2_00390 serine/threonine-protein kinase PpkA K11912 ppkA; serine/threonine-protein kinase PpkA [EC:2.7.11.1]
C PADK2_00425 hypothetical protein K11913 fha1; type VI secretion system protein
C PADK2_17390 transcriptional regulator K11914 sfa2; sigma-54 dependent transcriptional regulator
C PADK2_17355 serine/threonine phosphoprotein phosphatase Stp1 K11915 stp1; serine/threonine protein phosphatase Stp1 [EC:3.1.3.16]
C PADK2_00395 PppA K11915 stp1; serine/threonine protein phosphatase Stp1 [EC:3.1.3.16]
C PADK2_17350 serine-threonine kinase Stk1 K11916 stk1; serine/threonine-protein kinase Stk1 [EC:2.7.11.-]
C PADK2_13475 transcriptional regulator K11917 sfa3; sigma-54 dependent transcriptional regulator
C PADK2_13450 hypothetical protein K11918 lip3; type VI secretion system protein
C PADK2_13430 hypothetical protein K11919 hsiF3; type VI secretion system lysozyme-related protein
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AType VII secretion system
B ESX secretion system protein
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AChaperone-usher system
B Type 1 pilus assembly protein
C PADK2_03895 putative fimbrial subunit CupB1 K07345 fimA; major type 1 subunit fimbrin (pilin)
C PADK2_15035 fimbrial subunit CupA1 K07345 fimA; major type 1 subunit fimbrin (pilin)
C PADK2_20785 fimbrial subunit CupC1 K07345 fimA; major type 1 subunit fimbrin (pilin)
C PADK2_15030 chaperone CupA2 K07346 fimC; fimbrial chaperone protein
C PADK2_24710 pili assembly chaperone K07346 fimC; fimbrial chaperone protein
C PADK2_03905 usher CupB3 K07347 fimD; outer membrane usher protein
C PADK2_20775 usher CupC3 K07347 fimD; outer membrane usher protein
C PADK2_24715 hypothetical protein K07347 fimD; outer membrane usher protein
C PADK2_15025 usher CupA3 K07347 fimD; outer membrane usher protein
C PADK2_24695 hypothetical protein K27078 cupE; spore coat protein U domain-containing protein, fimbrial subunit CupE1/2/3/6
C PADK2_24700 hypothetical protein K27078 cupE; spore coat protein U domain-containing protein, fimbrial subunit CupE1/2/3/6
C PADK2_24720 hypothetical protein K27078 cupE; spore coat protein U domain-containing protein, fimbrial subunit CupE1/2/3/6
C PADK2_24705 hypothetical protein K27078 cupE; spore coat protein U domain-containing protein, fimbrial subunit CupE1/2/3/6
B P pilus assembly protein
B Mat/Ecp fimbriae assembly protein
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AExtracellular nucleation-precipitation pathway
B Curli assembly protein
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ASec (secretion) system
B Prokaryotic Sec-SRP core components
C PADK2_22945 secA; preprotein translocase subunit SecA K03070 secA; preprotein translocase subunit SecA [EC:7.4.2.8]
C PADK2_27135 preprotein translocase subunit SecB K03071 secB; preprotein translocase subunit SecB
C PADK2_05220 secD; preprotein translocase subunit SecD K03072 secD; preprotein translocase subunit SecD
C PADK2_03290 secE; preprotein translocase subunit SecE K03073 secE; preprotein translocase subunit SecE
C PADK2_05225 secF; preprotein translocase subunit SecF K03074 secF; preprotein translocase subunit SecF
C PADK2_25205 secG; preprotein translocase subunit SecG K03075 secG; preprotein translocase subunit SecG
C PADK2_03455 secY; preprotein translocase subunit SecY K03076 secY; preprotein translocase subunit SecY
C PADK2_05605 signal recognition particle protein K03106 SRP54; signal recognition particle subunit SRP54 [EC:3.6.5.4]
C PADK2_01875 signal recognition particle-docking protein FtsY K03110 ftsY; fused signal recognition particle receptor
C PADK2_05215 yajC; preprotein translocase subunit YajC K03210 yajC; preprotein translocase subunit YajC
C PADK2_29580 membrane protein insertase K03217 yidC; YidC/Oxa1 family membrane protein insertase
B Prokaryotic Sec-SRP related proteins
B Eukaryotic Sec-SRP protein
C PADK2_05605 signal recognition particle protein K03106 SRP54; signal recognition particle subunit SRP54 [EC:3.6.5.4]
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ATwin-arginine translocation (Tat) system
B Twin-arginine translocation (Tat) pathway protein
C PADK2_26835 tatA; twin arginine translocase protein A K03116 tatA; sec-independent protein translocase protein TatA
C PADK2_26840 tatB; sec-independent translocase K03117 tatB; sec-independent protein translocase protein TatB
C PADK2_26845 twin arginine-targeting protein translocase TatC K03118 tatC; sec-independent protein translocase protein TatC
!
#
#[ BRITE | KEGG2 | KEGG ]
#Last updated: December 11, 2023